2.6: Macroevolution
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Another word for macroevolution is speciation, the production of species, this is the level of evolution that Darwin studied, the kind that occurs over immense periods of time, where small changes accumulate to make life diverge into often drastically different forms. Now that we've introduced Mendel and cellular biology we can explain the two main pieces that Darwin was missing, heredity and variation.
Hox genes are a good example of one mechanism that enables drastic biological changes. Minor variations in the same small set of genes direct a zygote to grow into the shape of a worm, fly, mouse, or a human.
Exercise 2.6.1
“-patric” refers to geography. “allo” is Greek for other. Allopatric speciation happens when two populations are separated geographically; if you can't meet, you can't breed; in the absence of gene flow, variations accumulate through mutation and natural selection effects the two populations differently until they become separate species.
Darwin saw evolution as the gradual accumulation of changes, but looking closer at the fossil record, especially shellfish, later paleontologists began to question Darwin's ideas of phyletic gradualism, and proposed the theory of punctuated equilibrium, where the equilibrium of stasis or lack of change, is punctuated or broken by rapid evolutionary change. Remember that we are using the word "rapid" in a geological sense, usually it means hundreds of thousands of years.
Exercise 2.6.2
Skim this intro to punctuated equilibrium